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Global warming and human-driven impacts are changing the World’s ecological zones. This book applies the principles described in Bailey’s Ecosystem Geography: From Ecoregions to Sites, 2nd ed. (Springer 2009, 1st ed. 1996) to describe and characterize the major terrestrial and aquatic ecological zones of the Earth. Bailey’s system for classifying these zones has been adopted by major organizations such as the U.S. Forest Service and The Nature Conservancy and this book is a significant contribution to a long tradition of classifying and studying the world’s ecological regions or ecoregions. It includes two color maps that show the major ecoregions of the continents and oceans. Also included are: - 106 illustrations with 55 in full color - A new chapter on mountains is included. - There are new sections that address concerns about how eco regions are changing under the relentless influence of humans and climate change - Another new feature is the discussion of using eco regional patterns to transfer research results and select sites for detecting climate change effects on ecosystem distribution - Use of ecoregional patterns to design monitoring networks and sustainable landscapes - Fire regimes in different regional ecosystems and their management implications.

1 Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 1
1.1 Concepts of Ecosystem Regions, or Ecoregions . . . . . 1
1.2 Need for a Comparative System of Generic Regions . . . . 2
1.3 The Process of Defining . . . . . . . . . . . . . . . . . . . . . . . . . 4
1.4 The Role of Climate . . . . . . . . . . . . . . . . . . . . . . . . . . . 5
1.5 Macroclimate . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5
1.6 Advantages of the Ecoclimatic Approach . . . . . . . . . . . . 6
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 6
2 Oceanic Types and Their Controls . . . . . . . . . . . . . . . . . . . 9
2.1 Factors Controlling Ocean Hydrology . . . . . . . . . . . . 9
2.1.1 Latitude . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 9
2.1.2 Major Wind Systems . . . . . . . . . . . . . . . . . . . . . 9
2.1.3 Precipitation and Evaporation . . . . . . . . . . . . . . . 10
2.2 Types of Hydrologic Regions, or Oceanic Ecoregions . . . 12
2.2.1 Dietrich’s Oceanographic Classification . . . . . . . . 12
2.3 Distribution of the Oceanic Regions . . . . . . . . . . . . . . . . 15
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 16
3 Ecoregions of the Oceans . . . . . . . . . . . . . . . . . . . . . . . . . . . 17
3.1 500 Polar Domain . . . . . . . . . . . . . . . . . . . . . . . . . . . 17
3.1.1 510 Inner Polar Division . . . . . . . . . . . . . . . . . . . 17
3.1.2 520 Outer Polar Division . . . . . . . . . . . . . . . . . . 17
3.2 600 Temperate Domain . . . . . . . . . . . . . . . . . . . . . . . . . 18
3.2.1 610 Poleward Westerlies Division and 620
Equatorward Westerlies Division . . . . . . . . . . . . 19
3.2.2 630 Subtropical Division . . . . . . . . . . . . . . . . . . 20
3.2.3 640 High-Salinity Subtropical Division . . . . . . . . 20
3.2.4 650 Jet Stream Division . . . . . . . . . . . . . . . . . . . 21
3.2.5 660 Poleward Monsoon Division . . . . . . . . . . . . . 21
3.3 700 Tropical Domain . . . . . . . . . . . . . . . . . . . . . . . . . . 21
3.3.1 710 Tropical Monsoon Division . . . . . . . . . . . . . 21
3.3.2 720 High-Salinity Tropical Monsoon Division . . . 22
3.3.3 730 Poleward Trades Division . . . . . . . . . . . . . . 22
3.3.4 740 Trade Winds Division . . . . . . . . . . . . . . . . . 23
3.3.5 750 Equatorward Trades Division . . . . . . . . . . . . 23
3.3.6 760 Equatorial Countercurrent Division . . . . . . . . 24
3.4 Shelf . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 24
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 25
4 Continental Types and Their Controls . . . . . . . . . . . . . . . . 27
4.1 The Controls of Macroclimate . . . . . . . . . . . . . . . . . . . . 27
4.1.1 Latitude . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 27
4.1.2 Continental Position . . . . . . . . . . . . . . . . . . . . . . 28
4.1.3 Modified by Elevation . . . . . . . . . . . . . . . . . . . . 30
4.2 Types of Ecoclimatic Zones, or Continental Ecoregions . . . 33
4.2.1 Principles of Ecological Climate Regionalization . . . 33
4.2.2 The Ko¨ppen–Trewartha Classification
of Climates . . . . . . . . . . . . . . . . . . . . . . . . . . . . 36
4.3 Distribution of the Continental Regions . . . . . . . . . . . . . 38
4.4 Relationship to Other Ecoregional Zoning Systems . . . . . 40
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 43
5 Ecoregions of the Continents: The Polar Ecoregions . . . . . . 47
5.1 100 Polar Domain . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 47
5.2 110 Icecap Division . . . . . . . . . . . . . . . . . . . . . . . . . . . . 48
5.3 120 Tundra Division . . . . . . . . . . . . . . . . . . . . . . . . . . . 48
5.4 130 Subarctic Division . . . . . . . . . . . . . . . . . . . . . . . . . 51
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 54
6 The Humid Temperate Ecoregions . . . . . . . . . . . . . . . . . . . 55
6.1 200 Humid Temperate Domain . . . . . . . . . . . . . . . . . . . 55
6.2 210 Warm Continental Division . . . . . . . . . . . . . . . . . . . 56
6.3 220 Hot Continental Division . . . . . . . . . . . . . . . . . . . . . 59
6.4 230 Subtropical Division . . . . . . . . . . . . . . . . . . . . . . . . 60
6.5 240 Marine Division . . . . . . . . . . . . . . . . . . . . . . . . . . . 61
6.6 250 Prairie Division . . . . . . . . . . . . . . . . . . . . . . . . . . . 63
6.7 260 Mediterranean Division . . . . . . . . . . . . . . . . . . . . . . 66
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 68
7 The Dry Ecoregions . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 69
7.1 300 Dry Domain . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 69
7.2 310 Tropical/Subtropical Steppe Division . . . . . . . . . . . . 69
7.3 320 Tropical/Subtropical Desert Division . . . . . . . . . . . . 72
7.4 330 Temperate Steppe Division . . . . . . . . . . . . . . . . . . . 75
7.5 340 Temperate Desert Division . . . . . . . . . . . . . . . . . . . 77
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 79
8 The Humid Tropical Ecoregions . . . . . . . . . . . . . . . . . . . . . 81
8.1 400 Humid Tropical Domain . . . . . . . . . . . . . . . . . . . . . 81
8.2 410 Savanna Division . . . . . . . . . . . . . . . . . . . . . . . . . . 83
8.3 420 Rainforest Division . . . . . . . . . . . . . . . . . . . . . . . . . 84
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 87
9 The Mountain Ecoregions . . . . . . . . . . . . . . . . . . . . . . . . . . 89
9.1 M Mountains with Elevational Zonation . . . . . . . . . . . . . 89
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 94
10 Ecoregions and Climate Change . . . . . . . . . . . . . . . . . . . . . 95
10.1 Long-Term Climate Change . . . . . . . . . . . . . . . . . . 95
10.2 Use of the Ko¨ppen Climate Classification to Detect
Climate Change . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 97
10.3 Summary . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 102
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 102
11 Continental Patterns and Boundaries . . . . . . . . . . . . . . . . . 105
11.1 Pattern Within Zones . . . . . . . . . . . . . . . . . . . . . . . 105
11.2 Disturbance and Succession . . . . . . . . . . . . . . . . . . . . . 107
11.3 Boundaries Between Zones . . . . . . . . . . . . . . . . . . . . . 108
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 109
12 Applications of Ecoregional Patterns . . . . . . . . . . . . . . . . . . 111
12.1 Design for Sustainability . . . . . . . . . . . . . . . . . . . . . 111
12.1.1 Understand Ecosystem Pattern in Terms
of Process . . . . . . . . . . . . . . . . . . . . . . . . . . . 111
12.1.2 Use Pattern to Design Sustainable
Landscapes . . . . . . . . . . . . . . . . . . . . . . . . . . 111
12.2 Significance to Ecosystem Management . . . . . . . . . . . . 113
12.2.1 Local Systems Within Context . . . . . . . . . . . . 114
12.2.2 Spatial Transferability of Models . . . . . . . . . . 114
12.2.3 Links Between Terrestrial and Aquatic
Systems . . . . . . . . . . . . . . . . . . . . . . . . . . . . 115
12.2.4 Design of Sampling Networks . . . . . . . . . . . . 116
12.2.5 Transfer Information . . . . . . . . . . . . . . . . . . . 117
12.2.6 Determining Suitable Locations for Seed
Transfer . . . . . . . . . . . . . . . . . . . . . . . . . . . . 118
12.2.7 Understanding Landscape Fragmentation . . . . 119
12.2.8 Choosing Planting Strategies for Landscaping
and Restoration . . . . . . . . . . . . . . . . . . . . . . . 119
12.2.9 Environmental Risk Assessment . . . . . . . . . . . 119
12.2.10 Learn from Successful Ecological Designs and
Predict Establishment of Invasive Species . . . 120
12.2.11 Maintain and Restore Biodiversity . . . . . . . . . 120
12.2.12 Facilitate Conservation Planning . . . . . . . . . . 120
12.3 Significance to Research . . . . . . . . . . . . . . . . . . . . . . . 121
12.3.1 Restructuring Research Programs . . . . . . . . . . 122
12.3.2 Some Research Questions . . . . . . . . . . . . . . . . 122
12.3.3 Natural Ecosystem Patterns . . . . . . . . . . . . . . . 123
12.3.4 Effects of Climatic Variation . . . . . . . . . . . . . . 123
12.3.5 Relationships Between Vegetation and
Landform . . . . . . . . . . . . . . . . . . . . . . . . . . . . 123
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 124
13 Use of Fire Regimes at the Ecoregion Scale . . . . . . . . . . . . . 127
13.1 Ecosystem Structure and Process . . . . . . . . . . . . . . . 127
13.2 Range of Variation . . . . . . . . . . . . . . . . . . . . . . . . . . . 127
13.3 Different Ecoregions, Different Fire Regimes . . . . . . . . 128
13.3.1 Precolonial Fire Regimes (Vale) . . . . . . . . . . . 128
13.3.2 Fire Regime Types (The Nature Conservancy) . . . 128
13.3.3 Characterizing the US Wildfire Regimes
(Malamud et al.) . . . . . . . . . . . . . . . . . . . . . . . 130
13.3.4 Other Studies . . . . . . . . . . . . . . . . . . . . . . . . . 130
13.4 Use of Fire Regime at the Ecoregion Scale . . . . . . . . . . 132
13.5 Why Ecoregions Are Needed . . . . . . . . . . . . . . . . . . . . 134
13.6 Use of Ecosystem Patterns Within Ecoregions . . . . . . . 134
13.7 Future Range of Variation . . . . . . . . . . . . . . . . . . . . . . 134
References . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 135
14 Summary and Conclusions . . . . . . . . . . . . . . . . . . . . . . . . . 137
Appendix A: Air Masses and Frontal Zones . . . . . . . . . . . . . . . . 141
Appendix B: Common and Scientific Names . . . . . . . . . . . . . . . . 143
Appendix C: Conversion Factors . . . . . . . . . . . . . . . . . . . . . . . . 145
Appendix D: Comparison of Ecoregion and Related
Approaches . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 147
Glossary of Technical Terms . . . . . . . . . . . . . . . . . . . . . . . . . . . . 153
Bibliography . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 159
Maps . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 165
About the Author . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 171
Index . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 173
· · · · · · (收起)